Scotland's largest bird of prey, the white-tailed eagle, has bred in the east of the country for the first time in almost 200 years.
BBC Scotland has learned a pair of the birds have raised a male chick, after nesting at a secret location in Fife.
The country's last native white-tailed eagle was shot in Shetland in 1918.
The species only returned to the UK following a reintroduction project in the west of Scotland, which began on the Isle of Rum in 1975.
Environment Minister Paul Wheelhouse welcomed the news.
"I hope this will be the first of many of this magnificent species, which will eventually spread their territories right across Scotland," he said.
"I'd like to thank all the partners who have played their part in making this exciting and special event happen."