Some puffins discovered on beaches in France are believed to be from Skomer
Record numbers of seabirds have been killed in the recent storms on the Welsh coast, raising concerns some species are at risk.
The Wildlife Trust said many of the birds died from malnourishment because they found it difficult to find food during the storms.
An additional 11,000 puffins have perished in the storms in France.
The Wildlife Trust of South and West Wales said many birds have been washed up on Welsh beaches including Newgale, Broad Haven and Ferryside.
The trust, which manages the important seabird islands of Skomer and Skokholm on the Pembrokeshire coast, said birds had been badly hit by the storms.
"The birds have struggled to feed, probably because the sea is so rough," said Skomer warden Ed Stubbings.