Hundreds of puffins washed up on the east coast of the UK are likely to have died of starvation as a result of the recent severe weather.
RSPB Scotland said it has taken numerous calls from members of the public about the birds, found on beaches stretching from Aberdeen and Angus down to Northumberland.
It is the worst puffin "wreck" — the death of a large number of seabirds in a single incident — in almost 50 years, the conservation charity said.
Many razorbills and guillemots have also perished, prompting fears about the upcoming seabird breeding season.
Dr Barnaby Smith, from the Centre for Ecology & Hydrology which is part of the Natural Environment Research Council, said the birds may have been using up all their resources just fighting against the unseasonably cold temperatures and strong easterly winds.
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